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Exercises · 13.1

Q.Define growth, differentiation, development, dedifferentiation, redifferentiation, determinate growth, meristem and growth rate.

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These eight terms describe how a plant enlarges, matures its cells and unfolds its body plan through the coordinated processes of cell division, expansion and specialisation.

Growth is regarded as one of the most fundamental and conspicuous characteristics of a living being. It is defined as an irreversible and permanent increase in size of an organ, its parts, or even an individual cell. It is accompanied by metabolic processes (both anabolic and catabolic) that use energy, and it involves an increase in protoplasmic material seen as increase in fresh weight, dry weight, length, area, volume and cell number.

Differentiation is the process in which cells derived from the root and shoot apical meristems and cambium undergo structural changes in their cell walls and protoplasm to perform specific functions. For example, to form a tracheary element, a cell loses its protoplasm and develops a strong, lignified wall.

Development is a term that includes all the changes an organism goes through during its life cycle, from seed germination to senescence. Broadly, development = growth + differentiation, and it is under the control of both intrinsic (genetic, plant growth regulators) and extrinsic (light, temperature, water, oxygen, nutrition) factors.

Dedifferentiation is the phenomenon in which living differentiated cells, which had lost the capacity to divide, regain that capacity under certain conditions. Examples are the formation of interfascicular cambium and cork cambium from parenchyma cells.

Redifferentiation occurs when the dedifferentiated tissues again lose their capacity to divide and mature to perform specific functions, for example the secondary xylem and secondary cortex produced by the cambia.

Determinate growth is limited growth — growth that ceases once a structure attains a certain, characteristic size, as seen in leaves, flowers and fruits, in contrast to the indeterminate growth of roots and shoots.

A meristem is a localised group of actively and continuously dividing cells found at growing regions of the plant, such as the root and shoot apical meristems and the intercalary and lateral meristems (cambium).

Growth rate is the increased growth per unit time. It can be expressed mathematically, and growth may be arithmetic (only one daughter cell continues to divide) or geometric (both daughter cells continue to divide).

✓Final answer

Growth = permanent, irreversible increase in size; Differentiation = maturation of meristem-derived cells into specialised cells; Development = the sum of growth and differentiation across the life cycle; Dedifferentiation = mature cells regaining the power to divide; Redifferentiation = such cells again losing the power to divide and maturing; Determinate growth = limited growth reaching a fixed size; Meristem = a region of continuously dividing cells; Growth rate = increased growth per unit time.

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