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Botany · Ch 5 — Taxonomy and Systematic Botany

Family: Euphorbiaceae (Castor Family / Spurge Family)

5.13.4

Family: Euphorbiaceae (Castor Family / Spurge Family)

Euphorbiaceae (Castor/Spurge family; APG excludes Peraceae, Phyllanthaceae and Picrodendraceae from it) has 214 genera and about 5,600 species, well represented in Africa and South America. Habit: shrubs (Ricinus communis), trees (Emblica officinalis), herbs (Phyllanthus amarus), twiners (Tragia involucrata), or xerophytic cactus-like plants with flattened, succulent, leaf-like stems called phylloclades (Euphorbia) - usually with milky or watery latex. Leaf: stipulate or exstipulate, mostly simple, alternate, sometimes reduced/deciduous, palmately lobed (Ricinus) or deeply lobed (Manihot); stipules can modify into spines (E. splendens) or glandular hairs (Jatropha curcas). Inflorescence: highly variable - terminal raceme (Croton, Ricinus), catkin (Acalypha hispida), cyme (Jatropha), solitary axillary (Phyllanthus), or the family's unique cyathium (Euphorbia): a cup-like involucre of 4-5 fused sepaloid bracts holding a single central naked female flower (a tricarpellary gynoecium) surrounded by groups of naked male flowers (each a single jointed stamen), with a nectar gland on the involucre's rim - though it looks like one flower, a cyathium is actually an entire inflorescence. Flower: always unisexual, monoecious (Baliospermum) or dioecious (Bridelia), usually hypogynous. Perianth: tepals 0-5. Androecium: stamens ranging from 1 (Euphorbia) to numerous/branched (Ricinus), free or connate filaments. Gynoecium: tricarpellary (rarely bi-, tetra- or pentacarpellary), syncarpous, superior ovary, axile placentation, 3 styles each split into a feathery bilobed stigma. Fruit: capsule or schizocarp, often splitting violently into three one-seeded cocci called a regma (Ricinus). …

Table 5.16Systematic position of Euphorbiaceae
APG ClassificationBentham and Hooker Classification
Kingdom: PlantaeKingdom: Plantae
Clade: AngiospermsClass: Dicotyledonae
Clade: EudicotsSub-class: Monochlamydeae
Clade: RosidsSeries: Unisexuales
Figure 5.22Ricinus communis

What this figure shows. A labelled botanical plate of Ricinus communis (castor): the Habit of the tall shrub with palmately lobed leaves; the Male flower with a perianth of 5 tepals and numerous branched stamens crowded into stipitate phalanges (polyadelphous); the Female flower with a perianth of 3 tepals, a spiny trilobed ovary, three styles and bifid stigmas; separate floral diagrams for the male and female flowers; an L.S. and C.S. of the female flower showing the ovary, ovule and locule; and the floral formulae for the male flower (Br.,Eb …

Table 5.17Economic importance of the family Euphorbiaceae (selected examples)
Economic importanceBinomialUseful partUses
Food plantManihot esculenta (Maravalli kizhanku / Tapioca)Tuberous rootsRoots are rich in starch and used for preparing bread, biscuits, chips and other food stuffs
Oil plantRicinus communis (Castor)SeedsUsed as vegetable oil; ricinoleic acid in this oil eliminates acne-causing bacteria and acts as a laxative and lubricant
RubberHevea brasiliensis (Para rubber)Coagulated latexLatex is used in rubber products like tube and tyre