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Botany · Ch 9 — Tissue and Tissue System

Anatomy of Dicot Stem - Sunflower

9.7.3

Anatomy of Dicot Stem - Sunflower

A sunflower stem - the textbook's dicot stem example - again reads outside-in as a clear sequence, but with more internal zoning than the corresponding root. Its epidermis is a single layer of compactly-arranged, cutin-covered, rectangular parenchyma cells bearing numerous multicellular hairs, with chloroplasts normally absent. Its cortex is not homogeneous like the root's - it splits into three distinct functional zones: an outer, few-layered hypodermis of living, corner-thickened collenchyma cells giving mechanical strength to the still-elongating stem; a middle zone of chlorenchyma with conspicuous intercellular spaces (carrying out photosynthesis, and sometimes housing resin ducts); and an inner zone of ordinary parenchyma dedicated to food storage. The cortex's innermost layer is again an endodermis, but here it usually shows no true casparian strips - instead its starch-rich cells earn it the alternative name starch sheath, homologous to (but not identical in function to) the root's true endodermis. Everything inside the endodermis makes up the stele: a pericycle, and - arranged in a ring around a large central parenchymatous pith - the vascular bundles, an organisation given the specific name ** …

Figure 9.19T.S. of Dicot Stem - Sunflower

What this figure shows. A transverse section of a sunflower stem, from outside in: an epidermis with cuticle and multicellular hairs; a three-zoned cortex - collenchymatous hypodermis, chlorenchyma with scattered resin ducts, and parenchyma; a starch-sheath endodermis; a pericycle; and a eustele of open, collateral, conjoint vascular bundles (cambium present between xylem and phloem) arranged in a ring around a large central parenchymatous pith. NOTE ON RECOVERY: the source pages for this figure (pages 35-36, between the dicot-stem Pericycle description and Table 9.4) are two consecutive near-image-only pages - the single largest content gap in this chapter - so this cover is reconstructed from the Epidermis/Cortex/Stele/Pericycle prose either side of the gap plus Table 9.4's comparis …