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Botany · Ch 9 — Tissue and Tissue System

Parenchyma

9.2.1

Parenchyma

Parenchyma (Gk. para = beside, enchein = to pour) is the plant's default, all-purpose ground tissue - present in almost every organ, living, thin-walled and built of cellulose, with cells that can be oval, polyhedral, cylindrical, irregular, elongated or even armed with projections. Its defining visual feature is the prominent intercellular space typically left where several cells meet, which is what lets parenchyma double as a storage compartment (for water, air or 'ergastic' reserve substances) and still allow gas or fluid to move between cells; occasional cells that specialise in storing resin, tannins or calcium crystals are given the special name idioblasts. Because parenchyma is so functionally flexible, the book names five named sub-types built around a single job each: aerenchyma, whose huge intercellular air cavities give buoyancy and aeration to aquatic plants like Hydrilla and Nymphaea; storage parenchyma, packed with reserve food such as the starch of a root or stem tuber; stellate parenchyma, star-shaped with arms radiating outward to leave large air spaces, as in banana or Canna petioles; chlorenchyma, chloroplast-bearing parenchyma specialised for photosynthesis, a …

Figure 9.4Parenchyma

What this figure shows. A block of loosely-packed, thin-walled, roughly oval-to-polygonal parenchyma cells, drawn with clearly visible triangular intercellular spaces at the corners where three or more cells meet - the diagnostic featur …

Figure 9.5Types of Parenchyma: (a) Aerenchyma, (b) Storage parenchyma, (c) Stellate parenchyma, (d) Chlorenchyma, (e) Prosenchyma

What this figure shows. Five contrasting parenchyma cell types drawn side by side. (a) Aerenchyma - cells enclosing large air cavities/intercellular spaces, as in the floating stem of Hydrilla or Nymphaea, for aeration and buoyancy. (b) Storage parenchyma - cells packed with starch grains, as in a root/stem tuber. (c) Stellate (star-shaped) parenchyma with arms radiating from a central body, as in the petiole of Banana or Canna. (d) Chlorenchyma - chloroplast-filled cells shown as palisade (elongated, compact) and spongy (irregular, with air spaces) parenchyma of a leaf mesophyll. (e) Prosenchyma - parenchyma cells that have become elongated and pointed with slightly thickened walls, giving mechanical support. …