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Exercise 9.3 · Q9

Q.An important problem in fishery science is to estimate the number of fish presently spawning in streams and use this information to predict the number of mature fish or "recruits" that will return to the rivers during the reproductive period. If SS is the number of spawners and RR the number of recruits, the "Beverton-Holt spawner recruit function" is
[!FORMULA] R(S)=SαS+βR(S)=\dfrac{S}{\alpha S+\beta}
where α\alpha and β\beta are positive constants. Show that this function predicts approximately constant recruitment when the number of spawners is sufficiently large.

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"Approximately constant recruitment for large SS" means showing R(S)R(S) approaches a fixed finite value as S→∞S\to\infty — divide through by SS, the variable that is growing, to see this directly.

Step 1. Write down the model.

R(S)=SαS+β,α,β>0R(S)=\frac{S}{\alpha S+\beta},\qquad \alpha,\beta>0

Step 2. Divide numerator and denominator by SS (the highest power of SS present, valid for S>0S>0):

R(S)=S/S(αS+β)/S=1α+βSR(S)=\frac{S/S}{(\alpha S+\beta)/S}=\frac1{\alpha+\dfrac\beta S}

Step 3. Let S→∞S\to\infty. Since β\beta is a fixed positive constant, βS→0\dfrac\beta S\to0 as S→∞S\to\infty:

R(S) ⟶ 1α+0=1αR(S)\ \longrightarrow\ \frac1{\alpha+0}=\frac1\alpha …

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