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Q.Compare endosperm and perisperm. Describe the development of endosperm.

Bihar BsebBihar Board Intermediate 2026Subjective· 5mImportance★★★★★
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Endosperm (3n, from triple fusion) nourishes the embryo; perisperm (2n, leftover nucellus) is residual maternal tissue. Endosperm develops mostly by the free-nuclear method (repeated nuclear divisions, then wall formation).

Comparison of endosperm and perisperm:

Endosperm —

  • Formed by triple fusion (the second male gamete fuses with the two polar nuclei / secondary nucleus of the central cell) = double fertilisation.
  • It is triploid (3n).
  • It is a food-storing (nutritive) tissue that provides nourishment to the developing embryo.
  • It may be consumed during embryo development (non-endospermic/ex-albuminous seeds, e.g. pea, bean) or persist in the mature seed (endospermic/albuminous seeds, e.g. castor, maize, coconut).

Perisperm —

  • It is the persistent, unused/residual part of the nucellus that remains in some seeds.
  • It is diploid (2n) maternal (sporophytic) tissue.
  • It is usually dry and non-functional (not a product of fertilisation). Examples: black pepper, beet.

Development of endosperm:

After triple fusion the primary endosperm cell contains the primary endosperm nucleus (PEN, 3n). This divides to form the endosperm. There are three patterns:

  1. Free-nuclear type (most common) — The PEN undergoes repeated free-nuclear (mitotic) divisions without cell-wall formation, producing a large number of free nuclei in the common cytoplasm. Later, cell walls are laid down and it becomes cellular. (Coconut water is a free-nuclear endosperm and the surrounding white kernel is the cellular endosperm.) …

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